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"It was only in the context of this particular, functioning idiosyncratic whole that one could even say, in the first place, what the parts themselves were. " - so the opposite of synergy. is there a word for it? anyway, it does suggest novel compression methods.

"why this rather than that," - i think the standard answer in biology is path dependence (history). i am not sure you mean this to be one of your constraints.

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